Bosistoa pentacocca (F.Muell.) Baill.

Rutaceae

Baillon, H.E. (1873), Histoire des Plantes 4: 470, adnot.

Tree to 18 m, sometimes multi-stemmed. Bark grey to dark grey, blotchy, finely longitudinally fissured to scaly. Stems hairy and becoming hairless.

New growth pinkish red. Leaves opposite, compound, pinnate or trifoliate. Stipules absent. Petiole 1-8 cm long. Leaflets opposite, 3-13 leaflets per leaf, imparipinnate (terminal leaflet present). Lateral petiolule 1-15 mm long with swelling at base, terminal petiolule 5-40 mm long with swelling at junction with lamina. Leaflet blade narrow oblong, oblong-lanceolate, oblong-elliptic to oblong-obovate, elliptic, 4.5–27 cm long, 1–9 cm wide, base cuneate to obtuse, margin entire, toothed or crenate, apex acute to acuminate. Between 13–30 pairs of lateral veins. Oil dots very distinct. Surfaces hairless or rarely finely hairy.

Inflorescences terminal or axillary, paniculate. Flowers bisexual, actinomorphic, 5-merous. Sepals connate at base or up to nearly their full length, 0.7-1 mm long, triangular. Petals free, narrowly elliptic to oblanceolate, 3.5–4.5 mm long, spreading, white. Stamens 10, alternately unequal in length, filaments opposite petals shorter than filaments alternating with petals. Ovary of 5 (-rarely 4) more or less free carpels, the free carpels obscured by the woolly indumentum of the ovary, carpels share a common base and style, ovary superior, style laterally attached to each carpel.

Fruit dry consisting of 1 or 2 basally fused follicles, follicles 1.5–2.3 cm long, 1.5-3 cm wide, compressed, wrinkled, densely hairy, brown. Fruit outer layer (excocarp) woody, separate from inner layer (endocarp) which forms two parts, a membraneous to fleshy tissue more or less persistent on seed which tears away from the cartilaginous layer expelled from the fruit. Seed 1 per follicle, 1-2 cm long, subellipsoid or subreniform.

Features not available.

Occurs in CEQ, southwards to north eastern NSW. Found in subtropical and dry rainforest communities.

Two subspecies of Bosistoa pentacocca are recognised in Queensland (Forster 2013), Bosistoa pentacocca subsp. pentacocca and Bosistoa pentacocca subsp. connaricarpa. The former can be diagnosed by 3-5(-6) leaflet pairs; and 16-30 secondary veins per side of midrib; the latter by 1 or 2 (rarely 3) leaflet pairs; and 13-16 (-18) secondary veins per side of midrib. Bosistoa pentacocca subsp. pentacocca is endemic to SE Qld and NE NSW and Bosistoa pentacocca subsp. connaricarpa is distributed in a number of populations from NE of Gympie to Bowen. This profile information and associated coding has been adapted from Cooper & Cooper (2004), Forster (2013), Harden et al. (2014), and Hartley (2013).

Euodia pentacocca F.Muell. Fragmenta Phytographiae Australiae 3(19): 41, (1862). Type: "Ad flumina Richmond et Clarence River. Dr. Beckler." Bosistoa sapindiformis F.Muell. ex Benth. nom. illeg. Flora Australiensis 1: 359, (1863). Acradenia bosistoi F.Muell. nom. illeg. Monthly Notices of Papers and Proceedings of the Royal Society of Tasmania: 8, (1868). Bosistoa pentacocca (F.Muell.) Engl. nom. illeg. Die Naturlichen Pflanzenfamilien 4 (131-2): 114, fig. 64 J, (1896). Bosistoa connaricarpa Domin. Eighth Contribution to the Flora of Australia. Repertorium Specierum Novarum Regni Vegetabilis 12: 390, (1913). Type: "Queensland: Rockhampton, leg. Dallachy." Bosistoa pentacocca (F.Muell.) Baill. var. pentacocca.  Journal of the Arnold Arboretum 58(4): 423, (1977). Bosistoa pentacocca var. connaricarpa (Domin) T.G.Hartley. Journal of the Arnold Arboretum 58(4): 424, (1977). Bosistoa pentacocca var. dryanderensis T.G.Hartley. Journal of the Arnold Arboretum 58(4): 425, (1977). Type: "Queensland. NORTH KENNEDY DISTRICT: Mt. Dryander, … Moriarty 968 (CANB, holotype; BRI, isotype)".

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