Australian Tropical Rainforest Plants - Online edition
Embelia caulialata S.T.Reynolds
Reynolds, S.T. (1991) Austrobaileya 3(3): 362. Type: Queensland, Cape Tribulation, 16 05 S, 145 29 E, 28 May 1982, K.A.W.Williams 82050; holo: BRI.
Vine stem diameters to 9 cm recorded. Cambial layer turns purple on exposure. Pith often red on smaller stems.
Male flowers: Inflorescence a large panicle of racemes. Flowers perfumed, each flower about 4 mm diam. Calyx tube about 0.3 mm long, gland-dotted. Calyx lobes about 0.8 mm long, gland-dotted, margins fimbriate or laciniate. Petals gland-dotted, about 2 mm long, clothed in hairs or papillae. Staminal filaments about 0.5 mm long, anther connective marked by several rounded surface glands. Female flowers: Inflorescence a large hairy panicle. Flowers unpleasantly perfumed, about 3-4 mm diam. Sepals triangular, about 0.6-1 mm long, outer and inner surfaces clothed in hairs. Petals elliptic, about 2-3.5 x 1 mm, outer surface densely clothed in short hairs. Ovary +/- globose, about 1 mm high. Ovules two per ovary, embedded in the basal placenta.
Fruits globular, about 3.5-5.5 mm diam., calyx persistent at the base and style persistent at the apex. Placenta basal. Seeds globular, about 2-3 mm diam., outer surface sparsely dotted with dark resinous glands. Testa intruding slightly into the endosperm. Conspicuous yellow resin glands present in the endosperm. Embryo curved, transverse in the seed, about 1.5-2 x 0.2 mm. Cotyledons about 0.2 mm long, no wider than the radicle. Radicle straight or slightly curved.
Cotyledons ovate, about 7-12 x 4-9 mm, apex mucronate. Oil dots large, reddish or honey-coloured. Margins ciliate with about 3-7 teeth on each side. Lower surface clothed with gland-like hairs. First pair of leaves elliptical, margins serrate with about 6-9 teeth on each side. Lower surface of the leaf blades clothed in short, glandular hairs. Oil dots translucent, orbicular and elongated. At the tenth leaf stage: leaf blade elliptic to obovate, apex acuminate, base cuneate, margin with about 11-25 teeth on each side. Oil dots circular or elongated (cigar-shaped) pale or reddish in colour. Terminal bud clothed in short gland-like hairs and similarly the young stems which are also gland-dotted. Midrib depressed on the upper surface. Venation obscure. Stems narrowly 2-winged with a pair of wings extending from the point of attachment of each petiole down the stem to the next node. Seed germination time 41 to 78 days.